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I've done a little Googling and found a few discussions around the interwebs from guys in similar situations and they haven't had problems. Some suggest the o-rings are there to compensate for less than perfect mating surfaces between the cylinders and engine cases, but somehow I doubt that is their purpose.
Some guys run without a base gasket at all, to increase compression in an easy and free way (cheaper to run without a gasket than it is to deck the head).
So.... for my money I'd keep building Nothing is gonna break. Worst case is you might get a little weeping, but if you used fresh gaskets and do things right, I'd bet you'll be fine.
I was thinking about this too. The o-ring is a good seal, and will handle a little movement. So why the gasket?
I am guessing, but the gasket may be designed to keep dirt and liquids out of the o-ring groove. If the gasket isn't there, rain, water, dust, would accumulate in the area, eventually compromising the o-ring seal.
Another possibility is that the o-ring seals the cylinder sleeve, keeping oil and gases from getting between the steel cylinder sleeve and aluminum cylinder, while the gasket just seals the aluminum cylinder to the base.
You'll probably get away with not replacing it, but I would replace it if it was me....I think the o-ring is probably more important then the gasket...
There are plenty of bikes out there with aluminium cylinders and steel liners that only run a base gasket, I reckon you'll be fine, it was probably over engineered by Honda.
The 'O' rings prevent oil soaking up cylinder liner through capillary action.
Lots of Honda cylinders were not the best quality casting so oil could 'soak through' the alloy.
If your running stock bores it shouldn't be any problem at all.
I've done hundreds of 'big bore' motors with oversize liners, very few have had any oil weeping (Yamaha XS650 is a different matter as the base gasket isn't fully supported)
My 391cc CJ250 has very slight 'damp patch', but, you can see the liners in a couple of places.
It still doesn't actually leak or pool oil anywhere though (and I've given it some really sever beatings )
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Nothing is gonna break. Worst case is you might get a little weeping, but if you used fresh gaskets and do things right, I'd bet you'll be fine.
